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The list you are trying to post to is used as a sublist (a list fed with messages from another (ezmlm) list), but not properly set up as a sublist. Put the name of the parent list (‘origlist@orighost’) which exactly matches the SENDER of the original (or parent) list into DIR/sublist. Check the ownership of DIR/sublist, to make sure that the user controlling the list can read it.
Alternatively, use the ezmlm-make(1) -0 ‘origlist@orighost’ switch (see Customizing ezmlm-make operation).